Desmond enters the Animus black door, once again reliving the memories of his ancestor Ezio Auditore da Firenze, who is on his way to Masyaf desire to fulfill the dream of his father find Altair’s library, the assassin ancestor of Ezio and Desmond in the well of that changed forever the history of the assassins. Then, Subject 16 says that, if committed, can reach the funeral of Lucy Stillman, revealing to him that he killed at the end of the third part of the saga. He also noted that he, by the Animus, is a virus that must be removed, and will do everything possible to keep it active and aware than anyone of his presence. This tells your mind is shattered into three parts: the Altair, Ezio and that of his, and that can only despertar- if the colon is just visualize all the memories of Ezio and Altair, so the Animus has nothing more to show and thus divides the memories of everyone. Your mind is taken on an island, where the subject is 16.
Desmond is in a coma and is stuck in the Animus because his conscience is not destroyed.
The game begins shortly after the events in Brotherhood.
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